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PHOTOGRAPHY: REPTILIA: TESTUDINES

TESTUDINES
TURTLES & TORTOISES

The most recent checklist of living testudines* (Rhodin et al 2010) lists 328 species and 452 taxa (species and subspecies) in 97 genera. This paper also reports the extinction of 10 taxa (eight species and two subspecies). The testudines are divided between the Cryptodira (straight-necked turtles) and the Pleurodira (side-necked turtles) with the former containing eleven families and 77 genera, and the letter three families and 20 genera. The photo library contains representatives of 10 families, of cryptodirid turtles and all three families of pleurodirid turtles, with 50 genera represented by 88 species and 99 taxa (21.6% of all known taxa).

* Rhodin, A.G.J., P.P.vanDijk, J.B.Iverson & H.B.Shaffer 2010 Turtles of the World, 2010 Update: Annotated Checklist of Taxonomy, Synonymy, Distribution, and Conservation Status. Chelonian Research Foundation. This paper can be downloaded as a pdf from here.

Legend:
Photographic locality. Wherever possible precidence is given to wild specimens from known localitions, wherein the 'photographic locality' relates to the origin of the photographed specimens, rather than the taxon's entire range. The only exceptions are indicated by (captive) or (preserved specimen). These are either specimens photographed in captivity, where the locality given is a general distribution for the taxon, or preserved specimens photographed as biological voucher specimens.

Column D1. Digitized images - the bulk of the library consists of 35mm transparencies (mostly Kodachrome or Velvia) which are gradually being digitized [using Nikon CoolScan 4000 or Plustek OpticFilm 7600i] and stored on hard disc. 'X' indicates that digitized images are available, '[]' indicates that the transparencies of this taxon have yet to be digitized.

Column D2. Digital images - since 2005 digital images have also been taken [using Canon EOS 30D, 40D or 7D]. 'X' indicates that digital images are available in addition to digitized images for this taxon.

CRYPTODIRA
STRAIGHT-NECKED TURTLES

The Cryptodira or Straight-necked turtles contains eleven families, ten of which are represented in the photo library, only the Platysternidae being absent.

CARETTOCHELYIDAE
FLY RIVER TURTLE

The monotypic Carettochelyidae from New Guinea and northern Australia is represented in the photo library.

Carettochelyidae
Fly River turtle
FLY RIVER TURTLE
Carettochelys insculpta
Madang
Prov., Papua New Guinea
click on image to enlarge 
    Photographed specimen: D1. D2.
Carettochelys insculpta Fly River turtle PNG (Madang) X  

CHELONIIDAE
Sea Turtles

The Cheloniidae contains all the sea turtles except one, six species in six genera. Only two species are currently represented in the photo library.

Cheloniidae
Sea turtles
GREEN SEA TURTLE
Chelonia mydas
Rinca Is., Komodo Is., Indonesia
PACIFIC HAWKSBILL SEA TURTLE Eretmochelys imbricata bissa
Nagada Harbour, Madang Prov.,
Papua New Guinea
click on images to enlarge
    Photograph locality: D1. D2.
Chelonia mydas Green sea turtle PNG (Western), Indonesia (Komodo), Timor-Leste (Baucau) X X
Eretmochelys imbricata bissa Pacific hawksbill sea turtle PNG (Madang), Timor-Leste (Lautém) X X


CHELYDRIDAE
SNAPPING TURTLES

The Chelydridae from North America, contains two monotypic genera, both of which are represented in the photo library.

Chelydridae
Snapping turtles
COMMON SNAPPING TURTLE Chelydra serpentina serpentina
S outheast USA
ALLIGATOR SNAPPING TURTLE Macrochelys temmincki
Black Bayou, Louisiana, USA
click on images to enlarge
    Photograph locality: D1. D2.
Chelydra serpentina serpentina Common snapping turtle USA (captive) X  
Chelydra serpentina osceola Florida snapping turtle Florida X  
Macrochelys temmincki Alligator snapping turtle USA (Florida, Lousiana, Arkansas) X X

 


DERMATEMYDIDAE
CENTRAL AMERICAN RIVER TURTLE

The monotypic Dermatemydidae from Central America is represented in the photo library.

 

Dermatemydidae
C.American river turtle
CENTRAL AMERICAN RIVER TURTLE
Dermatemys mawii
Central America
click on image to enlarge 
    Photographed specimen: D1. D2.
Dermatemys mawii Central American river turtle Central America (captive) X  

 


DERMOCHELYIDAE
LEATHERBACK SEA TURTLE

The monotypic and widespread Dermochelyidae is represented in the photo library.

Dermochelyidae
Leatherback sea turtle
LEATHERBACK SEA TURTLE Dermochelys coriacea
(hatching) Grande Riviere, Trinidad
click on image to enlarge 
    Photographed specimen: D1. D2.
Dermochelys coriacea Leatherback sea turtle Trinidad X  



EMYDIDAE
AMERICAN POND, WOOD & BOX TURTLES

The American Emydidae contains two subfamilies: Deirochelyinae and Emydinae, which contains six genera with 39 and four genera with eleven species respectively. The photo library contains five genera and 12 species (14 taxa) and three genera with five species (8 taxa) respectively.

Deirochelyinae
Cooters & sliders
Emydinae
American wood & box turtles

EASTERN CUBAN SLIDER
Trachemys decussata decussata

Zapata, Bay of Pigs, Matanzas, Cuba

SPOTTED TURTLE
Clemmys guttata
Illinois, USA

ORNATE BOX TURTLE
Terrapene ornata ornata
Wisconsin, USA

click on images to enlarge
Deirochelyinae COOTERS & SLIDERS Photograph locality: D1. D2.
Chrysemys picta bellii Western painted turtle USA (Wisconsin) (captive) X  
Deirochelys reticularis miaria Western chicken turtle USA (Florida) (captive) X  
Graptemys barbouri Barbour's map turtle USA (Florida) X  
Graptemys nigrinoda nigrinoda Northern black-nobbed map turtle USA (Florida) (captive) X  
Graptemys pseudogeographica kohnii Mississippi map turtle USA (Louisiana) X  
Pseudemys concinna floridana Coastal plain cooter USA (Florida) X  
Pseudemys nelsoni Florida red-bellied cooter USA (Florida) (captive) X  
Pseudemys peninsularis Peninsula cooter USA (Florida) X  
Trachemys decussata decussata Eastern Cuban slider Cuba X  
Trachemys decussata angusta Western Cuban slider Cuba X  
Trachemys nebulosa nebulosa Baja California slider Mexico (Baja California Sur) X  
Trachemys scripta scripta Yellow-bellied slider USA (Florida) X  
Trachemys scripta elegans Red-eared slided USA (Florida, Louisiana), Costa Rica X  
Trachemys venusta venusta Meso-American slider Honduras (Colon) X  
Emydinae WOOD & BOX TURTLES      
Emys orbicularis fritzjuergenobsti Iberian pond turtle Spain (Madrid) X X
Emys orbicularis hellenica Greek pond turtle Greece (Thrace)   X
Clemmys guttata Spotted turtle USA (Illinois) (captive) X X
Glyptemys insculpta Wood turtle USA (New York) (captive) X  
Terrapene carolina bauri Florida box turtle USA (Florida) (captive) X  
Terrapene carolina yucatana Yucatan box turtle Mexico (Yucatan) (captive) X  
Terrapene nelsoni klauberi Northern spotted box turtle Mexico (Sonora) X  
Terrapene ornata ornata Ornate box turtle USA (Wisconsin) X  

Terrapene ornata luteola

Desert box turtle USA (Arizona) X X

 


GEOEMYDIDAE
ASIAN-AMERICAN POND, WOOD & BOX TURTLES

The Geoemydidae from Asia and also South America, containing 18 genera and 70 species, is represented in the photo library by eight genera and 13 species (14 taxa).

Geoemydidae
Asia-American pond, wood & box turtles

REEVES' TURTLE
Mauremys reevesi
Baucau, Baucau Distr., Timor-Leste

PALE ROOFED TURTLE
Pangshura smithii pallidiceps
Bardia National Park, Bheri, Nepal
EASTERN SPOTTED-LEGGED TURTLE
Rhinoclemmys punctularia punctularia Ilha Maracá, Roraima, Brazil
click on images to enlarge
    Photograph locality: D1. D2.
Batagur kachuga Red-crowned roofed turtle India (Madhya Pradesh) X  
Cuora aurocapitata Yellow-headed box turtle China (captive) X  
Cyclemys dentata Asian leaf turtle Thailand X  
Heosemys grandis Giant Asian pond turtle Southeast Asia (captive) X  
Mauremys reevesii Reeves' turtle Timor-Leste (Dili, Baucau) X X
Mauremys rivulata Balkan terrapin Greece (Thrace)   X
Melanochelys trijuga trijuga Indian black turtle India (Karnataka) X  
Melanochelys trijuga thermalis Sri Lankan black turtle Sri Lanka X  
Pangshura smithii pallidiceps Pale-footed roofed turtle Nepal X  
Pangshura tentoria circumdata Pink-ringed tent turtle India (Madhya Pradesh) X  
Rhinoclemmys annulata Brown wood turtle Honduras (Colon) X  
Rhinoclemmys diademata Maracaibo wood turtle Venezuela (captive) X  
Rhinoclemmys pulcherrima manni Central American wood turtle Central America (captive) X  
Rhinoclemmys punctularia punctularia Eastern spotted-legged turtle Brazil (Roraima), Guyana X  

 


KINOSTERNIDAE
MUD & MUSK TURTLES

The American Kinosternidae contains two subfamilies, although the Staurotyphinae is not represented in the photo library. The Kinosterninae, 22 species in two genera, is represented by one genus and five species (seven taxa).

Kinosterninae
Mud & musk turtles
SCORPION MUD TURTLE
Kinosternon scorpioides scorpioides

Santa Cruz do Arari, Ilha Marajo, Pará, Brazil
click on image to enlarge 
Kinosterninae MUD TURTLES Photograph locality: D1. D2.
Kinosternon alamosae Alamos mud turtle Mexico (Sonora) X  
Kinosternon integrum Mexican mud turtle Mexico (Sonora) X  
Kinosternon scorpioides scorpioides Scorpion mud turtle Brazil (Pará) X  
Kinosternon scorpioides albogulare White-throated mud turtle Honduras (Gracios a Dios), Costa Rica X  
Kinosternon sonoriensis Sonora mud turtle USA (Arizona) [] X
Kinosternon subrubrum subrubrum Eastern mud turtle USA (Florida) (captive) X  

Kinosternon subrubrum steindachneri

Florida mud turtle USA (Florida) X  

 


TESTUDINIDAE
TORTOISES

The Testudinidae contains approximately 19 genera and 58 species, and it is represented in the photo library by 13 genera and 24 species (26 taxa).

 

Testudinidae
Tortoises

ANGULATE TORTOISE
Chersina angulata
Jan Marais Reserve, W.Cape Prov., S.Africa

BURMESE BLACK GIANT TORTOISE
Manouria emys phayrei
Kaeng Krachen Nat. Park,
Phetcharburi Prov., Thailand
SPECKLED PADLOPER
Homopus signata
Springbok, N.Cape, S.Africa
click on images to enlarge
    Photograph locality: D1. D2.
Aldabrachelys dussumieri Aldabra giant tortoise Aldabra (captive) X  
Astrochelys yniphora Angonoka ploughshare tortoise Madagascar (captive) X  
Chelonoidis carbonaria Red-footed tortoise Brazil (Roraima), Venezuela (Apure) x  
Chelonoidis denticulata Yellow-footed tortoise Brazil (Roraima), Guyana x  
Chelonoidis nigra Charles Island giant tortoise Galapagos (captive) x  
Chersina angulata Angulate tortoise S.Africa (W.Cape) X  
Geochelone elegans Indian starred tortoise India (Orissa), Sri Lanka X  
Geochelone platynota Burmese star tortoise Myanmar (captive) X  
Geochelone sulcata African spurred tortoise Mauritania, S.Africa (W.Cape) X X
Gopherus agazzisii Desert tortoise USA (Arizona), Mexico (Sonora) X  
Gopherus berlandieri Texas tortoise USA (Texas) X  
Gopherus polyphemus Gopher tortoise USA (Florida) X  
Homopus areolatus Parrot -beaked padloper S.Africa (captive) X  
Homopus signatus Speckled padloper S.Africa (N.Cape) X  
Homopus solus Berger's Nama padloper S.Africa (captive) X  
Kinixys belliana belliana Bell's hingeback tortoise East Africa (captive) X  
Kinixys spekii Speke's hingeback tortoise Tanzania X  
Malacochersus tornieri Pancake tortoise East Africa (captive) X  
Manouria emys phayrei Burmese black giant tortoise Thailand X  
Stigmochelys pardalis Leopard tortoise Namibia x  

Eurotestudo hermanni hermanni

Western Hermann's tortoise W.Europe (captive) X  

Eurotestudo hermanni boettgeri

Eastern Hermann's tortoise Greece (Thrace) X X
Testudo gracea gracea Mediterranean spur-thighed tortoise Mediterranean (captive) X  
Testudo gracea ibera Greek spur-thighed tortoise Greece (Thrace)   X
Testudo horsfieldii horsfieldii Central Asian tortoise C.Asia (captive) x  
Testudo kleinmanni Egyptian tortoise N.Africa (captive) X  

 


TRIONYCHIDAE
SOFT-SHELLED TURTLES

The Trionychidae contains two subfamilies: Cyclanorbinae and Trionychinae, with three genera and six species, and eleven genera, with 24 species, respectively. These subfamilies are represented in the photo library by one genus with two species, and two genera with four species.

 

Cyclanorbinae
Flapneck turtles
Trionychinae
Softshell turtles
NORTHERN INDIAN FLAPNECK TURTLE
Lissemys punctata andersoni
Chambl River, Agra, Madhya Pradesh, India
MALAYAN SOFTSHELL TURTLE Dogania subplana
Long Naah, Baram River, Fourth Div., Sarawak, Borneo
click on images to enlarge
CYCLANORBINAE FLAPNECK TURTLES Photograph locality: D1. D2.
Lissemys punctata punctata Southern Indian flapneck turtle Sri Lanka (captive) X  
Lissemys punctata andersoni Northern Indian flapneck turtle India (Madhya Pradesh) X  
TRIONYCHINAE SOFTSHELL TURTLES      
Apalone ferox Florida softshell turtle USA (Florida) (captive) X  

Apalone mutica mutica

Midland smooth softshell turtle USA (captive) X  

Apalone spinifera pallida

Pallid spiny soft-shelled turtle

USA (Louisiana) X  
Dogania subplana Malayan softshell turtle Borneo (Sarawak) X  


PLEURODIRA
SIDE-NECKED TURTLES

The Pleurodira or Side-necked turtles occur in S.America, Africa and Australasia. Three families are recognised and all three are represented in the photo library.



CHELIDAE
AUSTRALO-AMERICAN SNAKE-NECKED & SIDE-NECKED TURTLES

The Chelidae is split (by some authors) into the American Chelinae and the Australasian Chelodinae is represented in the photo library. The former contains seven genera and 23 species while the latter contains eight genera and 31 species. They are represented in the photo library by four genera, and four species, and four genera, and eight species, respectively.

 

Chelinae
American side-necked turtles
Chelodinae
Australasian side & snake-necked turtles
GUYANAN SIDE-NECKED TURTLE Phrynops tuberosus
Orinduik, Guyana
TIMOR SNAKE-NECKED TURTLE Chelodina mccordi timorensis
Lake Ira Lalaro, Lautem Distr.,
Timor-Leste
click on images to enlarge
CHELINAE AMERICAN SIDE-NECKED TURTLES Photograph locality: D1. D2.
Chelus fimbriata Mata mata Brazil (Roraima), Venezuela (Apure), Guyana, Peru (Loreto) X  
Mesoclemmys gibba Gibba toad-headed turtle S.America (captive) X  
Phrynops tuberosus Guyanan side-necked turtle Guyana X  

Platemys platycephala platycephala

Eastern twist-necked turtle Brazil (Roraima) X  
CHELODINAE AUSTRALASIAN SIDE & SNAKE-NECKED TURTLES      

Chelodina mccordi timorensis

Timor snake-necked turtle

Timor-Leste (Lautém) X X
Chelodina pritchardi Kemp Welch River snake-necked turtle PNG (captive) X  
Elseya branderhorsti Southern New Guinea snapping turtle PNG (Western) X  
Elseya novaeguineae New Guinea snapping turtle PNG (Madang, Western) X  
Emydura macquarii Murray River short-necked turtle Australia (captive) X  
Emydura subglobosa Red-bellied short-neck turtle PNG (Western, Central) X  

Macrochelodina parkeri

Fly River snake-necked turtle

PNG (captive) X  

Macrochelodina rugosa

Northern snake-necked turtle

Australia (WA) (specimen) X  

 


PELOMEDUSIDAE
AFRICAN SIDE-NECKED TURTLES

The Pelomedusidae of Africa contains two genera and 19 species. It is represented by two species, one from each genus.

 

Pelomedusidae
American side-necked turtles
EAST AFRICAN BLACK MUD TURTLE
Pelusios subniger subniger
Muzei, Kasanga, Lake Tanganyika, Tanzania
click on image to enlarge 
    Photograph locality: D1. D2.

Pelomedusa subrufa subrufa

African helmeted turtle

Namibia X  

Pelusios subniger subniger

Eastern African black mud turtle

Tanzania X  

 


PODOCNEMIDIDAE
SOUTH AMERICAN RIVER TURTLES

The Podocnemidae of South America contains three genera and eight species, and is represented by two genera and four species.

 

Podocnemididae
S.American river turtles
YELLOW-SPOTTED AMAZON
SIDE-NECKED TURTLE
Podocnemis unifilis
Iquitos, Loreto, Peru
click on image to enlarge 
    Photograph locality: D1. D2.
Peltocephalus dumerilianus Big-headed Amazon river turtle S.America (captive) X  

Podocnemis expansa

Giant South America river turtle

Guyana, Brazil (Pará) X  
Podocnemis unifilis Yellow-spotted Amazon side-necked turtle Brazil (Roraima), Peru (Loreto) X  
Podocnemis vogli Llanos side-necked turtle Venezuela (Apure) []